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The gray wolf is the second most specialised member of the genus Canis, after the Ethiopian wolf, as demonstrated by its morphological adaptations to hunting large prey, its more gregarious nature, and its highly advanced expressive behavior. It is nonetheless closely related enough to smaller Canis species, such as the eastern wolf, coyote, and golden jackal.
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The evolution of the wolf occurred over a geologic time scale of 800,000 years, transforming the first Middle Pleistocene wolf specimen that is recognized as being morphologically similar to Canis lupus into today's dog, dingo and gray wolf. Ecological factors including habitat type, climate, prey specialization and predatory competition will greatly influence the wolf's genetic population structure and cranio-dental plasticity.
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